Modomics - A Database of RNA Modifications

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Name	Reaction type level 1	Reaction type level 2	Reaction type level 3	Input group	Output group	Introduced group name	Introduced group type	Atom address	Site	Modification level
A:Am	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	2′O	sugar	1
A:Ar(p)	phosphoribosylation	acylation	group addition	H	O[C@@H]1[C@H](OP(=O)(O)O)O[C@H](*)[C@H]1O	phosphoribosyl	phosphoribose	2′O	sugar	1
A:g6A	carbamoylation	acylation	group addition	H	*C(=O)NCC(=O)O	glycylcarbamoyl	acyl	N6	base/exocyclic	1
A:hn6A	carbamoylation	acylation	group addition	H	*C(=O)NC(C(O)=O)C(O)CC	hydroxynorvalylcarbamoyl	acyl	N6	base/exocyclic	1
A:I	deamination	NH2/O exchange	group exchange	NH2	*=O	oxo	oxo	C6	base/endocyclic	1
A:i6A	isopentenylation	alkylation	group addition	H	*CC=C(C)/C	isopenthenyl	hydrocarbon	N6	base/exocyclic	1
A:m1A	methylation	alkylation	group addition	none	*C	methyl	hydrocarbon	N1	base/endocyclic	1
A:m2A	methylation	alkylation	group addition	none	*C	methyl	hydrocarbon	C2	base/endocyclic	1
A:m6A	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	N6	base/exocyclic	1
A:m8A	methylation	alkylation	group addition	none	*C	methyl	hydrocarbon	C8	base/endocyclic	1
A:t6A	carbamoylation	acylation	group addition	H	*C(=O)NC(C(O)=O)C(O)C	threonylcarbamoyl	acyl	N6	base/exocyclic	1
ac4C:ac4Cm	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	2'O	sugar	2
acp3U:acp3Y	pseudouridine formation	isomerization	isomerization	none	none	none	none	N1, C5	base/endocyclic	2
Am:Im	deamination	NH2/O exchange	group exchange	NH2	*=O	oxo	oxo	C6	base/endocyclic	2
Am:m1Am	methylation	alkylation	group addition	none	*C	methyl	hydrocarbon	N1	base/endocyclic	2
Am:m6Am	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	N6	base/exocyclic	2
C:ac4C	acetylation	acylation	group addition	H	*C(=O)C	acetyl	acyl	N4	base/exocyclic	1
C:C+	agmatinylation	amination	group addition	O	*NCCCCCCNC(=N)N	agmatinyl	polyamine	C2	base/endocyclic	1
C:Cm	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	2′O	sugar	1
C:f5C	formylation	acylation	group addition	H	*C([H])=O	formyl	acyl	C5	base/endocyclic	3
C:hm5C	hydroxymethylation	hydroxymethylation	group addition	H	*CO	hydroxymethyl	hydroxyalkyl	C5	base/endocyclic	3
C:ho5C	hydroxylation	oxygen addition	group addition	H	*O	hydroxyl	hydroxyl	C5	base/endocyclic	1
C:k2C	lysylation	amination	group addition	O	*NCCCCC(N)C(O)=O	lysyl	amino acid	C2	base/endocyclic	1
C:m3C	methylation	alkylation	group addition	none	*C	methyl	hydrocarbon	N3	base/endocyclic	1
C:m4C	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	N4	base/exocyclic	1
C:m5C	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	C5	base/endocyclic	1
C:s2C	O/S exchange	O/S exchange	group exchange	O	*=S	thio	sulphur	C2	base/endocyclic	1
C:U	deamination	NH2/O exchange	group exchange	NH2	*=O	oxo	oxo	C4	base/endocyclic	0
chm5U:mchm5U	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	C5.O3	extra group	3
cm5U:chm5U	hydroxylation	oxygen addition	group addition	H	*O	hydroxyl	hydroxyl	C5.C1	extra group	2
cm5U:mcm5U	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	C5.O3	extra group	2
Cm:ac4Cm	acetylation	acylation	group addition	H	*C(=O)C	acetyl	acyl	N4	base/exocyclic	2
Cm:f5Cm	formylation	acylation	group addition	H	*C([H])=O	formyl	acyl	C5	base/endocyclic	4
Cm:m4Cm	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	N4	base/exocyclic	2
Cm:m5Cm	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	C5	base/endocyclic	2
cmnm5ges2U:nm5ges2U	decarboxymethylation	dealkylation	group removal	*C(=O)O	*[H]	hydrogen	hydrogen	C5.N2	extra group	4
cmnm5s2U:cmnm5ges2U	geranylation	alkylation	group addition	none	C\C(C)=C\CCC(\C)=C\C*	geranyl	hydrocarbon	C2.S1	extra group	3
cmnm5s2U:cmnm5se2U	S/Se exchange	S/Se exchange	group exchange	S	*[Se]	seleno	seleno	C2	base/endocyclic	3
cmnm5s2U:nm5s2U	decarboxymethylation	dealkylation	group removal	*C(=O)O	*[H]	hydrogen	hydrogen	C5.N2	extra group	3
cmnm5se2U:nm5se2U	decarboxymethylation	dealkylation	group removal	*C(=O)O	*[H]	hydrogen	hydrogen	C5.N2	extra group	4
cmnm5U:cmnm5s2U	O/S exchange	O/S exchange	group exchange	O	*=S	thio	sulphur	C2	base/endocyclic	2
cmnm5U:cmnm5Um	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	2′O	sugar	2
cmnm5U:nm5U	decarboxymethylation	dealkylation	group removal	*C(=O)O	*[H]	hydrogen	hydrogen	C5.N2	extra group	2
cmo5U:mcmo5U	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	C5.O4	extra group	4
CoApN:acCoApN	acetylation	acylation	group addition	H	*C(=O)C	acetyl	acyl	O5'.P3	extra group	2
CoApN:malonyl-CoApN	carboxylation	acylation	group addition	H	*C(=O)CC(=O)O	malonyl	acyl	O5'.P3	extra group	2
CoApN:succinyl-CoApN	carboxylation	acylation	group addition	H	*C(=O)CCC(=O)O	succinyl	acyl	O5'.P3	extra group	2
ct6A:t6A	hydration	hydration	group addition	none	O	water molecule	other	N6.O2, N6.C4	extra group	1
D:acp3D	3-amino-3-carboxypropylation	acylation	group addition	H	*CCC(N)C(=O)O	3-amino-3-carboxypropyl	amino acid	N3	base/endocyclic	2
D:m5D	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	C5	base/endocyclic	2
f5C:f5Cm	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	2′O	sugar	2
G:Gm	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	2′O	sugar	1
G:Gr(p)	phosphoribosylation	acylation	group addition	H	O[C@@H]1[C@H](OP(=O)(O)O)O[C@H](*)[C@H]1O	phosphoribosyl	phosphoribose	2′O	sugar	1
G:m1G	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	N1	base/endocyclic	1
G:m2G	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	N2	base/exocyclic	1
G:m7G	methylation	alkylation	group addition	none	*C	methyl	hydrocarbon	N7	base/endocyclic	1
ges2U:cmnm5ges2U	carboxymethylaminomethylation	aminoalkylation	group addition	H	*CNCC(=O)O	carboxymethylaminomethyl	aminoalkyl	C5	base/endocyclic	3
ges2U:nm5ges2U	aminomethylation	aminoalkylation	group addition	H	*CN	aminomethyl	aminoalkyl	C5	base/endocyclic	4
Gm:m1Gm	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	N1	base/endocyclic	2
Gm:m2Gm	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	N2	base/exocyclic	2
GpppN:m7GpppN	methylation	alkylation	group addition	none	*C	methyl	hydrocarbon	N7	base/endocyclic	3
hm5C:f5C	formylation	oxidation	group addition					None		3
hm5C:f5Cm	formylation	acylation	group addition		*C([H])=O	formyl	acyl	C5	base/endocyclic	4
hm6A:f6A	dehydrogenation	dehydrogenation	group removal	H, H	none	none	none	N6.C1, N6.O2	extra group	3
hn6A:ms2hn6A	methylthiolation	thioalkylation	group addition	H	*SC	methylthio	thioalkyl	C2	base/endocyclic	2
ho5U:cmo5U	carboxymethylation	alkylation	group addition	H	*CC(=O)O	carboxymethyl	carboxymethyl	C5.O1	extra group	3
ho5U:mo5U	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	C5.O1	extra group	2
ho5U:mo5U	methylation		group exchange		None	methyl group	methyl group		None	None
i6A:io6A	hydroxylation	oxygen addition	group addition	H	*O	hydroxyl	hydroxyl	C4.C5	extra group	2
i6A:ms2i6A	methylthiolation	thioalkylation	group addition	H	*SC	methylthio	thioalkyl	C2	base/endocyclic	2
I:Im	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	2′O	sugar	2
I:m1I	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	N1	base/endocyclic	2
Im:m1Im	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	N1	base/endocyclic	3
imG-14:imG	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	N3	base/endocyclic	3
imG-14:imG2	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	N1.C1	extra group	3
imG-14:yW-86	3-amino-3-carboxypropylation	acylation	group addition	H	*CCC(N)C(=O)O	3-amino-3-carboxypropyl	amino acid	N1.C1	extra group	3
imG2:mimG	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	N3	base/endocyclic	4
inm5U:inm5Um	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	2'O	sugar	4
io6A:ms2io6A	methylthiolation	thioalkylation	group addition	H	*SC	methylthio	thioalkyl	C2	base/endocyclic	3
m1A:A	demethylation	deoxygenation	group removal					None		0
m1A:m1Am	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	2′O	sugar	2
m1A:m1I	deamination	NH2/O exchange	group exchange	NH2	*=O	oxo	oxo	C6	base/endocyclic	2
m1G:imG-14	acetylation	condensation-cyclization	group addition	H,H2	=C(C)	anhydro-acetic acid	hydrocarbon	N1.C1, N2		2
m1G:m1Gm	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	2'O	sugar	2
m1I:m1Im	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	2′O	sugar	3
m1Y:m1acp3Y	3-amino-3-carboxypropylation	acylation	group addition	H	*CCC(N)C(=O)O	3-amino-3-carboxypropyl	amino acid	N3	base/endocyclic	3
m2,2G:m2,2Gm	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	2′O	sugar	3
m2,7G:m2,2,7G	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	N2	base/exocyclic	3
m2,7G:m2,7Gm	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	2′O	sugar	3
m2,7GpppN:m2,2,7GpppN	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	N2	base/exocyclic	5
m2A:m2,8A	methylation	alkylation	group addition	none	*C	methyl	hydrocarbon	C8	base/endocyclic	2
m2G:m2,2G	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	N2	base/exocyclic	2
m2G:m2Gm	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	2′O	sugar	2
m2Gm:m2,2Gm	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	N2	base/exocyclic	3
m2Gm:m2,7Gm	methylation	alkylation	group addition	none	*C	methyl	hydrocarbon	N7	base/endocyclic	3
m3C:C	demethylation	deoxygenation	group removal					C3		None
m3U:m3Um	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	2′O	sugar	2
m4,4C:m4,4Cm	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	2′O	sugar	3
m4C:m4,4C	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	N4	base/exocyclic	2
m4C:m4Cm	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	2′O	sugar	2
m4Cm:m4,4Cm	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	N4	base/exocyclic	3
m5C:hm5C	hydroxylation	hydroxylation	group addition		*CO	hydroxymethyl	hydroxyalkyl	None	base/endocyclic	2
m5C:m5Cm	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	2'O	sugar	2
m5U:m5s2U	O/S exchange	O/S exchange	group exchange	O	*=S	thio	sulphur	C2	base/endocyclic	2
m5U:m5Um	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	2′O	sugar	2
m6,6A:m6,6Am	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	2′O	sugar	3
m6A:A	demethylation	dealkylation	group removal	*C	*H	hydrogen	hydrogen	N6	base/exocyclic	0
m6A:hm6A	hydroxylation	oxygen addition	group addition	H	*O	hydroxyl	hydroxyl	N6.C1	extra group	2
m6A:m6,6A	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	N6	base/exocyclic	2
m6A:m6Am	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	2′O	sugar	2
m6A:ms2m6A	methylthiolation	thioalkylation	group addition	H	*SC	methylthio	thioalkyl	C2	base/endocyclic	2
m6Am:m6,6Am	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	N6	base/exocyclic	3
m7G:m2,7G	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	N2	base/exocyclic	2
m7GpppN:m2,7GpppN	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	N2	base/exocyclic	4
m8A:m2,8A	methylation	alkylation	group addition	none	*C	methyl	hydrocarbon	C2	base/endocyclic	2
mchm5U:mchm5Um	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	2'O	sugar	4
mcm5U:mchm5U	hydroxylation	oxygen addition	group addition	H	*O	hydroxyl	hydroxyl	C5.C1	extra group	3
mcm5U:mcm5s2U	O/S exchange	O/S exchange	group exchange	O	*=S	thio	sulphur	C2	base/endocyclic	3
mcm5U:mcm5Um	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	2'O	sugar	3
mcm5Um:mchm5Um	hydroxylation	oxygen addition	group addition	H	*O	hydroxyl	hydroxyl	C5.C1	extra group	4
mcmo5U:mcmo5Um	methylation	alkylation	group addition		*C	methyl	hydrocarbon	O5'.O3A	phosphate	3
mnm5s2U:mnm5ges2U	geranylation	alkylation	group addition	none	C\C(C)=C\CCC(\C)=C\C*	geranyl	hydrocarbon	C2.S1	extra group	5
mnm5s2U:mnm5se2U	S/Se exchange	S/Se exchange	group exchange	S	*[Se]	seleno	seleno	C2	base/endocyclic	5
mnm5U:mnm5s2U	O/S exchange	O/S exchange	group exchange	O	*=S	thio	sulphur	C2	base/endocyclic	4
mpN:mmpN	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	O5'.O3A	phosphate	3
ms2i6A:ms2io6A	hydroxylation	oxygen addition	group addition	H	*O	hydroxyl	hydroxyl	C4.C5	extra group	3
N:Nm	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	2ʹO	sugar	None
ncm5U:nchm5U	hydroxylation	oxygen addition	group addition	H	*O	hydroxyl	hydroxyl	C5.C1	extra group	2
ncm5U:ncm5s2U	O/S exchange	O/S exchange	group exchange	O	*=S	thio	sulphur	C2	base/endocyclic	2
ncm5U:ncm5Um	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	2′O	sugar	2
nm5ges2U:mnm5ges2U	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	C5.N2	extra group	5
nm5s2U:inm5s2U	isopentenylation	alkylation	group addition	H	*CC=C(C)/C	isopenthenyl	hydrocarbon	C5.N2	base/exocyclic	4
nm5s2U:mnm5s2U	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	C5.N2	extra group	4
nm5s2U:nm5ges2U	geranylation	alkylation	group addition	none	C\C(C)=C\CCC(\C)=C\C*	geranyl	hydrocarbon	C2.S1	extra group	4
nm5s2U:nm5se2U	S/Se exchange	S/Se exchange	group exchange	S	*[Se]	seleno	seleno	C2	base/endocyclic	4
nm5se2U:mnm5se2U	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	C5.N2	extra group	5
nm5U:inm5U	isopentenylation	alkylation	group addition	H	*CC=C(C)/C	isopenthenyl	hydrocarbon	C5.N2	base/exocyclic	3
nm5U:mnm5U	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	C5.N2	extra group	3
nm5U:nm5s2U	O/S exchange	O/S exchange	group exchange	O	*=S	thio	sulphur	C2	base/endocyclic	3
OHyW:o2yW	O-peroxidaton	oxygen addition	group addition	H	O	peroxide	peroxide	N1.O4	extra group	7
OHyWx:OHyWy	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	N1.O6	extra group	6
OHyWy:OHyW	methoxycarbonylation	acylation	group addition	H	C(=O)OC, C	methoxycarbonyl	methoxycarbonyl	N1.N5	extra group	7
oQ:Q	epoxide deoxygenation	deoxygenation	group removal	O	none	none	none	C7.C4,C7.C6	extra group	2
pN:mpN	methylation	alkylation	group addition	none	*C	methyl	hydrocarbon	O5'.O2A	phosphate	2
pN:pG(pN)	5ʹ-3ʹ guanylylation	nucleotide addition	group addition	OH	[O-]P(=O)(O)OC[C@@H]3O[C@H](n2cnc1c2N=C(N)NC1=O)[C@@H](O)[C@H]3O*	guanylyl	nucleotide	O5'.P3	phosphate	0
ppN:GpppN	5ʹ-5ʹ guanylylation	nucleotide addition	group addition	*P(=O)(O)OP(=O)(O)O	[O-]P(=O)(O)OC[C@@H]3O[C@H](n2cnc1c2N=C(N)NC1=O)[C@@H](O)[C@H]3O*	guanylyl	nucleotide	O5'.P3	phosphate	0
pppN:ApppN	group addition	group addition	group exchange		O	water molecule	other	None	extra group	0
pppN:AppppN	group addition	group addition	group addition		O	water molecule	other	None	extra group	1
pppN:ApppppN	group addition	group addition	group addition		*CC(=O)N	carbamoylmethyl	acyl	None	base/endocyclic	0
pppN:CoApN	group addition	group addition	group addition	*P(=O)(O)OP(=O)(O)O	SCCNC(=O)CCNC(=O)C(O)C(C)(C)COP([O-])(=O)OP([O-])(=O)OC[C@H]3O[C@@H](n2cnc1c(N)ncnc12)C(O)[C@H]3O*	dephospho-coenzyme A	other group	O5'.P3	phosphate	1
pppN:m6ApppN	group addition	group addition	group addition		O	water molecule	other	None	extra group	0
pppN:m6AppppN	group addition	group addition	group addition		O	water molecule	other	None	extra group	1
pppN:m6ApppppN	group addition	group addition	group addition		O	water molecule	other	None	extra group	1
pppN:m7GppppN	group addition	group addition	group addition		O	water molecule	other	None	extra group	1
pppN:mpppN	methylation	alkylation	group addition	none	*C	methyl	hydrocarbon	O5'.O2G	phosphate	1
pppN:NADpN	nicotinamide adenine dinucleotidylation	nucleotide addition	group addition	*P(=O)(O)OP(=O)(O)O	NC(=O)c1ccc[n+](c1)C5OC(COP([O-])(=O)OP([O-])(=O)OC[C@H]4O[C@@H](n3cnc2c(N)ncnc23)C(O)[C@H]4O*)C(O)C5O	nicotinamide adenine dinucleotide	nucleotide	O5'.P3	phosphate	1
pppN:pN	excision from a larger primary transcript	hydrolysis	group removal	*P(=O)(O)OP(=O)(O)O	*[H]	hydrogen	hydrogen	O5'.O3A	phosphate	0
pppN:ppN	dephosphorylation	hydrolysis	group removal	*P(O)(O)=O	*[H]	hydrogen	hydrogen	O5'.O3B	phosphate	1
preQ0:G+	amination	amination	group addition	none	N, [H]	ammonia	ammonia	C7.C1, C7.N2	extra group	1
preQ0base:preQ0	transglycosylation	base exchange	group exchange	NC2=Nc1ncnc1C(=O)N2	NC2=Nc1n(*)cc(C#N)c1C(=O)N2	preQ0 base	nucleobase	C1ʹ	sugar	0
preQ0base:preQ1base	hydrogenation	hydrogenation	group addition	none	*H	hydrogen	hydrogen	C7.N2	extra group	-1
preQ1:oQ	2,3-epoxy-4,5-dihydroxycyclopentanylation	alkylation	group addition	H	*C2C1OC1C(O)C2O	2,3-epoxy-4,5-dihydroxycyclopentane group	epoxycyclopentane group	C7.N2	extra group	1
preQ1base:preQ1	transglycosylation	base exchange	group exchange	NC2=Nc1ncnc1C(=O)N2	NC2=Nc1ncc(CN)c1C(=O)N2	preQ1 base	nucleobase	C1ʹ	sugar	0
Q:galQ	galactosylation	glycosylation	group addition	H	O[C@@H]1[C@@H](O)[C@@H](O)[C@@H](CO)O[C@@]1(O)*	galactosyl	saccharide	C7.O5	extra group	3
Q:gluQ	glutamylation	acylation	group addition	H	*OC(=O)C(N)CCC(=O)O	glutamyl	amino acid	C7.O5	base/exocyclic	3
Q:manQ	mannosylation	glycosylation	group addition	H	OC1[C@H](O)[C@H](O)C(CO)O[C@]1(*)[H]	mannosyl	saccharide	C7.O5	extra group	3
Q:Qbase	N-glycosyl bond hydrolysis	hydrolysis	group removal	C1=CC(C(C1NCC2=CN(C3=C2C(=O)N=C(N3)N)C4C(C(C(O4)CO)O)O)O)O	*O	hydroxyl	hydroxyl	C1ʹ	sugar	0
Qbase:Q	transglycosylation	base exchange	group exchange	NC2=Nc1ncnc1C(=O)N2	C1=CC(C(C1NCC2=CN(C3=C2C(=O)N=C(N3)N)C4C(C(C(O4)CO)O)O)O)O	Qbase	nucleobase	C1ʹ	sugar	0
s2U:cmnm5s2U	carboxymethylaminomethylation	aminoalkylation	group addition	H	*CNCC(=O)O	carboxymethylaminomethyl	aminoalkyl	C5	base/endocyclic	2
s2U:ges2U	geranylation	alkylation	group addition	none	C\C(C)=C\CCC(\C)=C\C*	geranyl	hydrocarbon	C2.S1	extra group	2
s2U:nm5s2U	aminomethylation	aminoalkylation	group addition	H	*CN	aminomethyl	aminoalkyl	C5	base/endocyclic	3
s2U:s2Um	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	2'O	sugar	2
s2U:se2U	S/Se exchange	S/Se exchange	group exchange	S	*[Se]	seleno	seleno	C2	base/endocyclic	2
s2U:tm5s2U	taurinomethylation	aminoalkylation	group addition	H	*CNCCS(=O)(O)=O	taurinomethyl	aminoalkyl	C5	base/endocyclic	2
se2U:cmnm5se2U	carboxymethylaminomethylation	aminoalkylation	group addition	H	*CNCC(=O)O	carboxymethylaminomethyl	aminoalkyl	C5	base/endocyclic	3
se2U:nm5se2U	aminomethylation	aminoalkylation	group addition	H	*CN	aminomethyl	aminoalkyl	C5	base/endocyclic	4
t6A:ct6A	dehydration	dehydration	group removal	H, *O	none	none	none	N6.O2, N6.C4	extra group	2
t6A:m6t6A	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	N6	base/exocyclic	2
t6A:ms2t6A	methylthiolation	thioalkylation	group addition	H	*SC	methylthio	thioalkyl	C2	base/endocyclic	2
tm5U:tm5s2U	O/S exchange	O/S exchange	group exchange	O	*=S	thio	sulphur	C2	base/endocyclic	2
U:acp3U	3-amino-3-carboxypropylation	acylation	group addition	H	*CCC(N)C(=O)O	3-amino-3-carboxypropyl	amino acid	N3	base/endocyclic	1
U:cm5U	carboxymethylation	alkylation	group addition	H	*CC(=O)O	carboxymethyl	carboxymethyl	C5	base/endocyclic	1
U:cmnm5U	carboxymethylaminomethylation	aminoalkylation	group addition	H	*CNCC(=O)O	carboxymethylaminomethyl	aminoalkyl	C5	base/endocyclic	1
U:D	hydrogenation	hydrogenation	group addition	none	H, H	hydrogen	hydrogen	C5,C6	base/endocyclic	1
U:ho5U	hydroxylation	oxygen addition	group addition	H	*O	hydroxyl	hydroxyl	C5	base/endocyclic	1
U:m3U	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	N3	base/endocyclic	1
U:m5U	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	C5	base/endocyclic	1
U:mcm5s2U	thiolation		group addition					None		1
U:ncm5U	carbamoylmethylation	alkylation	group addition	H	*CC(=O)N	carbamoylmethyl	acyl	C5	base/endocyclic	1
U:nm5U	aminomethylation	aminoalkylation	group addition	H	*CN	aminomethyl	aminoalkyl	C5	base/endocyclic	2
U:s2U	O/S exchange	O/S exchange	group exchange	O	*=S	thio	sulphur	C2	base/endocyclic	1
U:s4U	O/S exchange	O/S exchange	group exchange	O	*=S	thio	sulphur	C4	base/endocyclic	1
U:tm5U	taurinomethylation	aminoalkylation	group addition	H	*CNCCS(=O)(O)=O	taurinomethyl	aminoalkyl	C5	base/endocyclic	1
U:Um	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	2′O	sugar	1
U:Y	pseudouridine formation	isomerization	isomerization	none	none	none	none	N1, C5	base/endocyclic	1
Um:cmnm5Um	carboxymethylaminomethylation	aminoalkylation	group addition	H	*CNCC(=O)O	carboxymethylaminomethyl	aminoalkyl	C5	base/endocyclic	2
Um:s2Um	O/S exchange	O/S exchange	group exchange	O	*=S	thio	sulphur	C2	base/endocyclic	2
Y:acp3Y	3-amino-3-carboxypropylation	acylation	group addition	H	*CCC(N)C(=O)O	3-amino-3-carboxypropyl	amino acid	N3	base/endocyclic	2
Y:m1Y	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	N1	base/endocyclic	2
Y:m3Y	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	N3	base/endocyclic	2
Y:Ym	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	2′O	sugar	2
yW-58:yW	methoxycarbonylation	acylation	group addition	H	*C(=O)OC	methoxycarbonyl	methoxycarbonyl	N1.N5	extra group	6
yW-72:OHyWx	hydroxylation	oxygen addition	group addition	H	*O	hydroxyl	hydroxyl	N1.C3	extra group	5
yW-72:yW-58	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	N1.C6	extra group	5
yW-86:yW-72	methylation	alkylation	group addition	H	*C	methyl	hydrocarbon	N3	base/endocyclic	4


A:Am
A:Ar(p)
A:g6A
A:hn6A
A:I
A:i6A
A:m1A
A:m2A
A:m6A
A:m8A
A:t6A
ac4C:ac4Cm
acp3U:acp3Y
Am:Im
Am:m1Am
Am:m6Am
C:ac4C
C:C+
C:Cm
C:f5C
C:hm5C
C:ho5C
C:k2C
C:m3C
C:m4C
C:m5C
C:s2C
C:U
chm5U:mchm5U
cm5U:chm5U
cm5U:mcm5U
Cm:ac4Cm
Cm:f5Cm
Cm:m4Cm
Cm:m5Cm
cmnm5ges2U:nm5ges2U
cmnm5s2U:cmnm5ges2U
cmnm5s2U:cmnm5se2U
cmnm5s2U:nm5s2U
cmnm5se2U:nm5se2U
cmnm5U:cmnm5s2U
cmnm5U:cmnm5Um
cmnm5U:nm5U
cmo5U:mcmo5U
CoApN:acCoApN
CoApN:malonyl-CoApN
CoApN:succinyl-CoApN
ct6A:t6A
D:acp3D
D:m5D
f5C:f5Cm
G:Gm
G:Gr(p)
G:m1G
G:m2G
G:m7G
ges2U:cmnm5ges2U
ges2U:nm5ges2U
Gm:m1Gm
Gm:m2Gm
GpppN:m7GpppN
hm5C:f5C
hm5C:f5Cm
hm6A:f6A
hn6A:ms2hn6A
ho5U:cmo5U
ho5U:mo5U
ho5U:mo5U
i6A:io6A
i6A:ms2i6A
I:Im
I:m1I
Im:m1Im
imG-14:imG
imG-14:imG2
imG-14:yW-86
imG2:mimG
inm5U:inm5Um
io6A:ms2io6A
m1A:A
m1A:m1Am
m1A:m1I
m1G:imG-14
m1G:m1Gm
m1I:m1Im
m1Y:m1acp3Y
m2,2G:m2,2Gm
m2,7G:m2,2,7G
m2,7G:m2,7Gm
m2,7GpppN:m2,2,7GpppN
m2A:m2,8A
m2G:m2,2G
m2G:m2Gm
m2Gm:m2,2Gm
m2Gm:m2,7Gm
m3C:C
m3U:m3Um
m4,4C:m4,4Cm
m4C:m4,4C
m4C:m4Cm
m4Cm:m4,4Cm
m5C:hm5C
m5C:m5Cm
m5U:m5s2U
m5U:m5Um
m6,6A:m6,6Am
m6A:A
m6A:hm6A
m6A:m6,6A
m6A:m6Am
m6A:ms2m6A
m6Am:m6,6Am
m7G:m2,7G
m7GpppN:m2,7GpppN
m8A:m2,8A
mchm5U:mchm5Um
mcm5U:mchm5U
mcm5U:mcm5s2U
mcm5U:mcm5Um
mcm5Um:mchm5Um
mcmo5U:mcmo5Um
mnm5s2U:mnm5ges2U
mnm5s2U:mnm5se2U
mnm5U:mnm5s2U
mpN:mmpN
ms2i6A:ms2io6A
N:Nm
ncm5U:nchm5U
ncm5U:ncm5s2U
ncm5U:ncm5Um
nm5ges2U:mnm5ges2U
nm5s2U:inm5s2U
nm5s2U:mnm5s2U
nm5s2U:nm5ges2U
nm5s2U:nm5se2U
nm5se2U:mnm5se2U
nm5U:inm5U
nm5U:mnm5U
nm5U:nm5s2U
OHyW:o2yW
OHyWx:OHyWy
OHyWy:OHyW
oQ:Q
pN:mpN
pN:pG(pN)
ppN:GpppN
pppN:ApppN
pppN:AppppN
pppN:ApppppN
pppN:CoApN
pppN:m6ApppN
pppN:m6AppppN
pppN:m6ApppppN
pppN:m7GppppN
pppN:mpppN
pppN:NADpN
pppN:pN
pppN:ppN
preQ0:G+
preQ0base:preQ0
preQ0base:preQ1base
preQ1:oQ
preQ1base:preQ1
Q:galQ
Q:gluQ
Q:manQ
Q:Qbase
Qbase:Q
s2U:cmnm5s2U
s2U:ges2U
s2U:nm5s2U
s2U:s2Um
s2U:se2U
s2U:tm5s2U
se2U:cmnm5se2U
se2U:nm5se2U
t6A:ct6A
t6A:m6t6A
t6A:ms2t6A
tm5U:tm5s2U
U:acp3U
U:cm5U
U:cmnm5U
U:D
U:ho5U
U:m3U
U:m5U
U:mcm5s2U
U:ncm5U
U:nm5U
U:s2U
U:s4U
U:tm5U
U:Um
U:Y
Um:cmnm5Um
Um:s2Um
Y:acp3Y
Y:m1Y
Y:m3Y
Y:Ym
yW-58:yW
yW-72:OHyWx
yW-72:yW-58
yW-86:yW-72

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