ID MODOMICS code Name Short Name RNAMods code Moiety type Formula Monoisotopic mass Average mass Degeneracy Nature in DNA in RNA Non-modified counterpart(s) Nucleotide RCSB ligand code Related nucleotide MODOMICS code Related nucleoside MODOMICS code
340 nucleotide None None Natural None None Adenosine 32
341 nucleotide None None Natural None None Guanosine 63
342 nucleotide None None Natural None None Adenosine 33
343 nucleotide None None Natural None None Uridine 8
344 nucleotide None None Natural None None Adenosine 26
345 nucleotide None None Natural None None Uridine 88
346 nucleotide None None Natural None None Cytidine 101
347 nucleotide None None Natural None None Cytidine 92
348 nucleotide None None Natural None None Uridine 138
349 nucleotide None None Natural None None Uridine 48
350 nucleotide None None Natural None None Cytidine 2
351 nucleotide None None Natural None None Uridine 85
352 nucleotide None None Natural None None Uridine 145
353 nucleotide None None Natural None None Uridine 146
354 nucleotide None None Natural None None Uridine 144
355 nucleotide None None Natural None None Uridine 115
356 nucleotide None None Natural None None Uridine 43
357 nucleotide None None Natural None None Uridine 73
358 nucleotide None None Natural None None Uridine 87
359 nucleotide None None Natural None None Uridine 44
360 nucleotide None None Natural None None Uridine 64
361 nucleotide None None Natural None None Uridine 80
362 nucleotide None None Natural None None Uridine 27
363 nucleotide None None Natural None None Uridine 74
364 nucleotide None None Natural None None Uridine 17
365 nucleotide None None Natural None None Guanosine 59
366 nucleotide None None Natural None None Cytidine 183
367 nucleotide None None Natural None None Adenosine 128
368 nucleotide None None Natural None None Uridine 103
369 nucleotide None None Natural None None Uridine 179
370 nucleotide None None Natural None None Guanosine 34
371 nucleotide None None Natural None None Adenosine 136
372 nucleotide None None Natural None None Adenosine 182
373 nucleotide None None Natural None None Uridine 166
374 nucleotide None None Natural None None Adenosine 180
375 nucleotide None None Natural None None Adenosine 126
376 nucleotide None None Natural None None Adenosine 79
377 nucleotide None None Natural None None Adenosine 5
378 nucleotide None None Natural None None Adenosine 55
379 nucleotide None None Natural None None Uridine 117
380 nucleotide None None Natural None None Uridine 57
381 nucleotide None None Natural None None Uridine 137
382 nucleotide None None Natural None None Uridine 75
383 nucleotide None None Natural None None Guanosine 108
384 nucleotide None None Natural None None Uridine 164
385 nucleotide None None Natural None None Uridine 169
386 nucleotide None None Natural None None Uridine 62
387 nucleotide None None Natural None None Uridine 35
388 nucleotide None None Natural None None Uridine 163
389 nucleotide None None Natural None None Uridine 29
390 nucleotide None None Natural None None Uridine 162
391 nucleotide None None Natural None None Uridine 11
392 nucleotide None None Natural None None Uridine 23
393 nucleotide None None Natural None None Uridine 167
394 nucleotide None None Natural None None Uridine 54
395 nucleotide None None Natural None None Uridine 45
396 nucleotide None None Natural None None Uridine 178
397 nucleotide None None Natural None None Cytidine 104
398 nucleotide None None Natural None None Cytidine 143
399 nucleotide None None Natural None None Uridine 3
400 nucleotide None None Natural None None Uridine 168
401 nucleotide None None Natural None None Uridine 78
402 nucleotide None None Natural None None Uridine 110
403 nucleotide None None Natural None None Uridine 91
404 nucleotide None None Natural None None Uridine 21
405 nucleotide None None Natural None None Guanosine 71
406 nucleotide None None Natural None None Guanosine 77
407 nucleotide None None Natural None None Guanosine 112
408 nucleotide None None Natural None None Guanosine 123
409 nucleotide None None Natural None None Guanosine 120
410 nucleotide None None Natural None None Adenosine 131
411 nucleotide None None Natural None None Cytidine 141
412 nucleotide None None Natural None None Guanosine 99
413 nucleotide None None Natural None None Adenosine 175
414 nucleotide None None Natural None None Guanosine 6
415 nucleotide None None Natural None None Guanosine 86
416 nucleotide None None Natural None None Guanosine 42
417 nucleotide None None Natural None None Adenosine 181
418 nucleotide None None Natural None None Guanosine 97
419 nucleotide None None Natural None None Guanosine 89
420 nucleotide None None Natural None None Guanosine 90
421 nucleotide None None Natural None None Guanosine 160
422 nucleotide None None Natural None None Guanosine 121
423 nucleotide None None Natural None None Guanosine 65
424 nucleotide None None Natural None None Guanosine 93
425 nucleotide None None Natural None None Uridine 58
426 nucleotide None None Natural None None Guanosine 139
427 nucleotide None None Natural None None Cytidine 69
428 nucleotide None None Natural None None Guanosine 102
429 nucleotide None None Natural None None Guanosine 40
430 nucleotide None None Natural None None Adenosine 82
431 nucleotide None None Natural None None Adenosine 124
432 nucleotide None None Natural None None Adenosine 10
433 nucleotide None None Natural None None Adenosine 125
434 nucleotide None None Natural None None Adenosine 177
435 nucleotide None None Natural None None Adenosine 114
436 nucleotide None None Natural None None Adenosine 176
437 nucleotide None None Natural None None Adenosine 19
438 nucleotide None None Natural None None Adenosine 130
439 nucleotide None None Natural None None Adenosine 130
440 nucleotide None None Natural None None Guanosine 4
441 nucleotide None None Natural None None Guanosine 111
297 511551U (2R,4S)-1-[(4R)-3,4-dihydroxytetrahydrofuran-2-YL]-5-[(methylamino)methyl]-1,2,3,4-tetrahydropyrimidine-2,4-diol-5'-monophosphate pmnm5U nucleotide None 367.249 not aplicable Natural No Yes Uridine

MNU

129
32 01A 1,2'-O-dimethyladenosine m1Am œ nucleoside C12O4N5H17 295.1281 295.294 not applicable Natural No Yes Adenosine
63 01G 1,2'-O-dimethylguanosine m1Gm ε nucleoside C12O5N5H17 311.123 311.294 not applicable Natural No Yes Guanosine
33 019A 1,2'-O-dimethylinosine m1Im ξ nucleoside C12O5N4H16 296.1121 296.279 not applicable Natural No Yes Adenosine
230 20551U 1-(5-O-phosphono-beta-D-ribofuranosyl)-2-selanylpyrimidin-4(1H)-one pse2U nucleotide None 387.142 not aplicable Natural No Yes Uridine

RUS

94
187 2551U 1-(beta-D-ribofuranosyl)-2-thio-uracil-5'-phosphate ps2U nucleotide None 340.247 not aplicable Natural No Yes Uridine

SUR

100
8 1309U 1-methyl-3-(3-amino-3-carboxypropyl)pseudouridine m1acp3Y α nucleoside C14O8N3H21 359.1329 359.332 not applicable Natural No Yes Uridine
83 1A 1-methyladenosine m1A " nucleoside C11O4N5H15 281.1124 281.268 not applicable Natural No Yes Adenosine 204
22 1G 1-methylguanosine m1G K nucleoside C11O5N5H15 297.1073 297.267 not applicable Natural No Yes Guanosine 214
26 19A 1-methylinosine m1I O nucleoside C11O5N4H14 282.0964 282.253 not applicable Natural No Yes Adenosine
88 19U 1-methylpseudouridine m1Y ] nucleoside C10O6N2H14 258.0852 258.228 not applicable Natural No Yes Uridine
214 1551G 1N-methylguanosine-5'-monophosphate pm1G nucleotide None 377.247 not aplicable Natural No Yes Guanosine

1MG

22
296 05551U 2',5-dimethyluridine-5'-monophosphate pm5Um nucleotide None 352.234 not aplicable Natural No Yes Uridine

2MU

122
256 00551A 2'-O-[(5'-phospho)ribosyl]adenosine-5'-monophosphate pAr(p) nucleotide None 559.316 not aplicable Natural No Yes Adenosine

RIA

20
183 051C 2'-O-methyl-5-hydroxymethylcytidine hm5Cm ¡ nucleoside C11H17N3O6 287.1117 287.269 not applicable Natural No Yes Cytidine
127 0A 2'-O-methyladenosine Am : nucleoside C11O4N5H15 281.1124 281.268 not applicable Natural No Yes Adenosine 207
207 0551A 2'-O-methyladenosine 5'-(dihydrogen phosphate) pAm nucleotide None 361.248 not aplicable Natural No Yes Adenosine

A2M

127
84 0C 2'-O-methylcytidine Cm B nucleoside C10O5N3H15 257.1012 257.243 not applicable Natural No Yes Cytidine 201
28 0G 2'-O-methylguanosine Gm # nucleoside C11O5N5H15 297.1073 297.267 not applicable Natural No Yes Guanosine 193
128 09A 2'-O-methylinosine Im nucleoside C11O5N4H14 282.0964 282.253 not applicable Natural No Yes Adenosine
103 09U 2'-O-methylpseudouridine Ym Z nucleoside C10O6N2H14 258.0852 258.228 not applicable Natural No Yes Uridine
50 0U 2'-O-methyluridine Um J nucleoside C10O6N2H14 258.0852 258.228 not applicable Natural No Yes Uridine 205
179 0503U 2'-O-methyluridine 5-oxyacetic acid methyl ester mcmo5Um Ͽ nucleoside C13H18N2O9 346.1012 346.29 not applicable Natural No Yes Uridine
20 00A 2'-O-ribosyladenosine (phosphate) Ar(p) ^ nucleoside C15O11N5H22P1 479.1053 479.336 not applicable Natural No Yes Adenosine 256
34 00G 2'-O-ribosylguanosine (phosphate) Gr(p) nucleoside C15O12N5H22P1 495.1003 495.335 not applicable Natural No Yes Guanosine
174 3377N 2'3'-cyclic phosphate end N2'3'cp nucleotide None 272.043 unspecified residue Natural No Yes Adenosine, Uridine, Cytidine, Guanosine
136 28A 2,8-dimethyladenosine m2,8A ± nucleoside C12H17N5O4 295.1281 295.294 not applicable Natural No Yes Adenosine
182 21161A 2- methylthiomethylenethio-N6-isopentenyl-adenosine msms2i6A £ nucleoside C17H25N5O4S2 427.1347 427.541 not applicable Natural No Yes Adenosine
166 21U 2-geranylthiouridine ges2U Γ nucleoside C19H28N2O5S 396.1719 396.501 not applicable Natural No Yes Uridine
101 21C 2-lysidine k2C } nucleoside C15O6N5H25 371.1805 371.389 not applicable Natural No Yes Cytidine
37 2A 2-methyladenosine m2A / nucleoside C11O4N5H15 281.1124 281.268 not applicable Natural No Yes Adenosine 219
219 2551A 2-methyladenosine-5'-monophosphate pm2A nucleotide None 361.248 not aplicable Natural No Yes Adenosine

2MA

37
180 2164A 2-methylthio cyclic N6-threonylcarbamoyladenosine ms2ct6A ÿ nucleoside C17H21N5O7S 439.1162 439.443 not applicable Natural No Yes Adenosine
126 2160A 2-methylthio-N6-(cis-hydroxyisopentenyl) adenosine ms2io6A nucleoside C16O5N5H23S1 397.142 397.449 not applicable Natural No Yes Adenosine
79 2163A 2-methylthio-N6-hydroxynorvalylcarbamoyladenosine ms2hn6A nucleoside C17O8N6H24S1 472.1376 472.473 not applicable Natural No Yes Adenosine
240 2161551A 2-methylthio-N6-isopentenyl-adenosine-5'-monophosphate pms2i6A nucleotide None 461.43 not aplicable Natural No Yes Adenosine

MIA

51
51 2161A 2-methylthio-N6-isopentenyladenosine ms2i6A * nucleoside C16O4N5H23S1 381.1471 381.45 not applicable Natural No Yes Adenosine 240
5 621A 2-methylthio-N6-methyladenosine ms2m6A nucleoside C12O4N5H17S1 327.1001 327.359 not applicable Natural No Yes Adenosine
55 2162A 2-methylthio-N6-threonylcarbamoyladenosine ms2t6A [ nucleoside C16O8N6H22S1 458.122 458.446 not applicable Natural No Yes Adenosine
94 20U 2-selenouridine se2U ω nucleoside C9O5N2H12Se1 305.9919 307.162 not applicable Natural No Yes Uridine 230
117 02U 2-thio-2'-O-methyluridine s2Um nucleoside C10O5N2H14S1 274.0623 274.294 not applicable Natural No Yes Uridine
92 2C 2-thiocytidine s2C % nucleoside C9O4N3H13S1 259.0627 259.282 not applicable Natural No Yes Cytidine
100 2U 2-thiouridine s2U 2 nucleoside C9O5N2H12S1 260.0467 260.267 not applicable Natural No Yes Uridine 187
191 2551G 2N-methylguanosine-5'-monophosphate pm2G nucleotide None 377.247 not aplicable Natural No Yes Guanosine

2MG

36
57 03U 3,2'-O-dimethyluridine m3Um σ nucleoside C11O6N2H16 272.1008 272.255 not applicable Natural No Yes Uridine
137 308U 3-(3-amino-3-carboxypropyl)-5,6-dihydrouridine acp3D Ð nucleoside C13H21N3O8 347.1329 347.321 not applicable Natural No Yes Uridine
138 309U 3-(3-amino-3-carboxypropyl)pseudouridine acp3Y Þ nucleoside C13H19N3O8 345.1172 345.305 not applicable Natural No Yes Uridine
75 30U 3-(3-amino-3-carboxypropyl)uridine acp3U X nucleoside C13O8N3H19 345.1172 345.305 not applicable Natural No Yes Uridine
299 3551C 3-Methylcytidine- 5'-monophosphate pm3C nucleotide None 337.223 not aplicable Natural No Yes Cytidine

JMH

52
52 3C 3-methylcytidine m3C ' nucleoside C10O5N3H15 257.1012 258.251 not applicable Natural No Yes Cytidine 299
48 39U 3-methylpseudouridine m3Y κ nucleoside C10O6N2H14 258.0852 258.228 not applicable Natural No Yes Uridine
49 3U 3-methyluridine m3U δ nucleoside C10O6N2H14 258.0852 258.228 not applicable Natural No Yes Uridine 199
199 3551U 3-methyluridine-5'-monophosphate pm3U nucleotide None 338.208 not aplicable Natural No Yes Uridine

UR3

49
108 4G 4-demethylwyosine imG-14 nucleoside C13O5N5H15 321.1073 321.289 not applicable Natural No Yes Guanosine
281 4551C 4-methyl, cytidine-5'-monophosphate pm4C nucleotide None 337.223 not aplicable Natural No Yes Cytidine

B8T

9
7 74U 4-thiouridine s4U 4 nucleoside C9O5N2H12S1 260.0467 260.267 not applicable Natural No Yes Uridine 216
216 74551U 4-thiouridine-5'-monophosphate ps4U nucleotide None 340.247 not aplicable Natural No Yes Uridine

4SU

7
208 04551C 4N,O2'-methylcytidine-5'-monophosphate pm4Cm nucleotide None 351.25 not aplicable Natural No Yes Cytidine

4OC

39
149 455N 5' (3' -dephospho-CoA) CoApN nucleotide C21H32N7O13P2S 684.1254 879.618 unspecified residue Natural No Yes Adenosine, Uridine, Cytidine, Guanosine
150 4155N 5' (3' -dephosphoacetyl-CoA) acCoApN nucleotide C22H40N7O15P2S 736.1778 921.654 unspecified residue Natural No Yes Adenosine, Uridine, Cytidine, Guanosine
151 4255N 5' (3' -dephosphomalonyl-CoA) malonyl-CoApN nucleotide C23H40N7O17P2S 780.1676 965.664 unspecified residue Natural No Yes Adenosine, Uridine, Cytidine, Guanosine
152 4355N 5' (3' -dephosphosuccinyl-CoA) succinyl-CoApN nucleotide C24H41N7O17P2S 793.1755 979.69 unspecified residue Natural No Yes Adenosine, Uridine, Cytidine, Guanosine
153 552N 5' diphosphate end ppN ϒ nucleotide PO3 78.9585 291.066 unspecified residue Natural No Yes Adenosine, Uridine, Cytidine, Guanosine
173 550N 5' hydroxyl end 5'-OH-N nucleoside None None unspecified residue Natural No Yes Adenosine, Uridine, Cytidine, Guanosine
154 255N 5' nicotinamide adenine dinucleotide NADpN Ξ nucleotide C21H25N7O14P2 661.0935 856.496 unspecified residue Natural No Yes Adenosine, Uridine, Cytidine, Guanosine
147 553N 5' triphosphate end pppN ϖ nucleotide P3O9R 157.917 369.031 unspecified residue Natural No Yes Adenosine, Uridine, Cytidine, Guanosine
2 05C 5,2'-O-dimethylcytidine m5Cm τ nucleoside C11O5N3H17 271.1168 271.27 not applicable Natural No Yes Cytidine
122 05U 5,2'-O-dimethyluridine m5Um \ nucleoside C11O6N2H16 272.1008 272.255 not applicable Natural No Yes Uridine 296
190 8551U 5,6-dihydrouridine-5'-monophosphate pD nucleotide None 326.197 not aplicable Natural No Yes Uridine

H2U

14
223 2521551U 5-(O-methylaceto)-2-thio-2-deoxy-uridine-5'-monophosphate pmcm5s2U nucleotide None 412.31 not aplicable Natural No Yes Uridine

70U

68
164 0522U 5-(carboxyhydroxymethyl)-2'-O-methyluridine methyl ester mchm5Um b nucleoside C13H18N2O9 346.1012 346.29 not applicable Natural No Yes Uridine
85 522U 5-(carboxyhydroxymethyl)uridine methyl ester mchm5U , nucleoside C12O9N2H16 332.0856 332.263 not applicable Natural No Yes Uridine
241 502551U 5-(carboxymethoxy) uridine-5'-monophosphate pcmo5U nucleotide None 398.217 not aplicable Natural No Yes Uridine

CM0

47
254 51551C 5-(hydroxymethyl)cytidine 5'-(dihydrogen phosphate) phm5C nucleotide None 353.223 not aplicable Natural No Yes Cytidine

5HM

119
145 0583U 5-(isopentenylaminomethyl)-2'-O-methyluridine inm5Um ¼ nucleoside C16H25N3O6 355.1743 355.386 not applicable Natural No Yes Uridine
146 2583U 5-(isopentenylaminomethyl)-2-thiouridine inm5s2U ½ nucleoside C15H23N3O5S 357.1358 357.425 not applicable Natural No Yes Uridine
144 583U 5-(isopentenylaminomethyl)uridine inm5U ¾ nucleoside C15H23N3O6 341.1587 341.36 not applicable Natural No Yes Uridine
169 21510U 5-aminomethyl-2-geranylthiouridine nm5ges2U Δ nucleoside C20H31N3O5S 425.1984 425.542 not applicable Natural No Yes Uridine
62 20510U 5-aminomethyl-2-selenouridine nm5se2U π nucleoside C10O5N3H15Se1 337.0176 336.203 not applicable Natural No Yes Uridine
35 2510U 5-aminomethyl-2-thiouridine nm5s2U nucleoside C10O5N3H15S1 289.0732 289.308 not applicable Natural No Yes Uridine
115 510U 5-aminomethyluridine nm5U nucleoside C10O6N3H15 273.0961 273.243 not applicable Natural No Yes Uridine
163 531U 5-carbamoylhydroxymethyluridine nchm5U r nucleoside C11H15N3O8 317.0859 317.252 not applicable Natural No Yes Uridine
29 053U 5-carbamoylmethyl-2'-O-methyluridine ncm5Um ~ nucleoside C12O7N3H17 315.1066 315.279 not applicable Natural No Yes Uridine
162 253U 5-carbamoylmethyl-2-thiouridine ncm5s2U l nucleoside C11H15N3O6S 317.0681 317.318 not applicable Natural No Yes Uridine
43 53U 5-carbamoylmethyluridine ncm5U & nucleoside C11O7N3H15 301.091 301.253 not applicable Natural No Yes Uridine
73 520U 5-carboxyhydroxymethyluridine chm5U nucleoside C11O9N2H14 318.0699 318.237 not applicable Natural No Yes Uridine
11 2540U 5-carboxymethyl-2-thiouridine cm5s2U nucleoside C11O7N2H14S1 318.0522 318.303 not applicable Natural No Yes Uridine
23 051U 5-carboxymethylaminomethyl-2'-O-methyluridine cmnm5Um ) nucleoside C13O8N3H19 345.1172 345.305 not applicable Natural No Yes Uridine
167 2151U 5-carboxymethylaminomethyl-2-geranylthiouridine cmnm5ges2U f nucleoside C22H33N3O7S 483.2039 483.578 not applicable Natural No Yes Uridine
54 2051U 5-carboxymethylaminomethyl-2-selenouridine cmnm5se2U nucleoside C12O7N3H17Se1 393.0239 394.239 not applicable Natural No Yes Uridine
45 251U 5-carboxymethylaminomethyl-2-thiouridine cmnm5s2U $ nucleoside C12O7N3H17S1 347.0787 347.344 not applicable Natural No Yes Uridine
87 51U 5-carboxymethylaminomethyluridine cmnm5U ! nucleoside C12O8N3H17 331.1016 331.279 not applicable Natural No Yes Uridine
44 52U 5-carboxymethyluridine cm5U nucleoside C11O8N2H14 302.075 302.237 not applicable Natural No Yes Uridine
178 55U 5-cyanomethyluridine cnm5U Ѷ nucleoside C11H13N3O6 283.0804 283.237 not applicable Natural No Yes Uridine
104 071C 5-formyl-2'-O-methylcytidine f5Cm ° nucleoside C11O6N3H15 285.0961 285.253 not applicable Natural No Yes Cytidine
31 71C 5-formylcytidine f5C > nucleoside C10O6N3H13 271.0804 271.227 not applicable Natural No Yes Cytidine 232
232 71551C 5-formylcytidine 5'-(dihydrogen phosphate) pf5C nucleotide None 351.207 not aplicable Natural No Yes Cytidine

RSQ

31
143 50C 5-hydroxycytidine ho5C Ç nucleoside C9H13N3O6 259.0804 259.216 not applicable Natural No Yes Cytidine
119 51C 5-hydroxymethylcytidine hm5C nucleoside C10O6N3H15 273.0961 273.243 not applicable Natural No Yes Cytidine 254
64 50U 5-hydroxyuridine ho5U nucleoside C9O7N2H12 260.0645 260.201 not applicable Natural No Yes Uridine
80 0521U 5-methoxycarbonylmethyl-2'-O-methyluridine mcm5Um nucleoside C13O8N2H18 330.1063 330.291 not applicable Natural No Yes Uridine
68 2521U 5-methoxycarbonylmethyl-2-thiouridine mcm5s2U 3 nucleoside C12O7N2H16S1 332.0678 332.33 not applicable Natural No Yes Uridine 223
27 521U 5-methoxycarbonylmethyluridine mcm5U 1 nucleoside C12O8N2H16 316.0907 316.264 not applicable Natural No Yes Uridine
74 501U 5-methoxyuridine mo5U 5 nucleoside C10O7N2H14 274.0801 274.227 not applicable Natural No Yes Uridine
3 25U 5-methyl-2-thiouridine m5s2U F nucleoside C10O5N2H14S1 274.0623 274.294 not applicable Natural No Yes Uridine
168 21511U 5-methylaminomethyl-2-geranylthiouridine mnm5ges2U h nucleoside C21H33N3O5S 439.2141 439.569 not applicable Natural No Yes Uridine
78 20511U 5-methylaminomethyl-2-selenouridine mnm5se2U nucleoside C11O5N3H17Se1 349.0341 350.23 not applicable Natural No Yes Uridine
110 2511U 5-methylaminomethyl-2-thiouridine mnm5s2U S nucleoside C11O5N3H17S1 303.0889 303.335 not applicable Natural No Yes Uridine
129 511U 5-methylaminomethyluridine mnm5U { nucleoside C11O6N3H17 287.1117 287.269 not applicable Natural No Yes Uridine 297
18 5C 5-methylcytidine m5C ? nucleoside C10O5N3H15 257.1012 257.243 not applicable Natural No Yes Cytidine 186
186 5551C 5-methylcytidine-5'-monophosphate pm5C nucleotide None 337.223 not aplicable Natural No Yes Cytidine

5MC

18
17 58U 5-methyldihydrouridine m5D ρ nucleoside C10O6N2H16 260.1008 260.244 not applicable Natural No Yes Uridine
12 5U 5-methyluridine m5U T nucleoside C10O6N2H14 258.0852 260.244 not applicable Natural No Yes Uridine 194
194 5551U 5-methyluridine-5'-monophosphate pm5U nucleotide None 338.208 not aplicable Natural No Yes Uridine

5MU

12
91 254U 5-taurinomethyl-2-thiouridine tm5s2U nucleoside C12O8N3H19S2 397.0614 397.425 not applicable Natural No Yes Uridine
21 54U 5-taurinomethyluridine tm5U Ê nucleoside C12O9N3H19S1 381.0842 381.359 not applicable Natural No Yes Uridine
204 1551A 6-hydro-1-methyladenosine-5'-monophosphate pm1A nucleotide None 361.248 not aplicable Natural No Yes Adenosine

1MA

83
192 66551A 6N-dimethyladenosine-5'-monophosphate pm6,6A nucleotide None 375.274 not aplicable Natural No Yes Adenosine

MA6

41
71 47G 7-aminocarboxypropyl-demethylwyosine yW-86 ¥ nucleoside C17O7N6H22 422.155 422.393 not applicable Natural No Yes Guanosine
77 347G 7-aminocarboxypropylwyosine yW-72 Ω nucleoside C18O7N6H24 436.1706 436.419 not applicable Natural No Yes Guanosine
112 348G 7-aminocarboxypropylwyosine methyl ester yW-58 nucleoside C19O7N6H26 450.1863 450.446 not applicable Natural No Yes Guanosine
61 101000G 7-aminomethyl-7-carbaguanine preQ1base base C7O1N5H9 179.0807 179.179 not applicable Natural No Yes Guanosine 123
123 101G 7-aminomethyl-7-deazaguanosine preQ1 nucleoside C12O5N5H17 311.123 311.294 not applicable Natural No Yes Guanosine
95 100000G 7-cyano-7-carbaguanine preQ0base ψ base C7O1N5H5 175.0494 175.148 not applicable Natural No Yes Guanosine 120
120 100G 7-cyano-7-deazaguanosine preQ0 φ nucleoside C12O5N5H13 307.0917 307.262 not applicable Natural No Yes Guanosine
116 7G 7-methylguanosine m7G 7 nucleoside C11O5N5H17 299.123 298.275 not applicable Natural No Yes Guanosine 203
203 7551G 7N-methyl-8-hydroguanosine-5'-monophosphate pm7G nucleotide None 379.263 not aplicable Natural No Yes Guanosine

7MG

116
131 8A 8-methyladenosine m8A â nucleoside C11O4N5H15 281.1124 281.268 not applicable Natural No Yes Adenosine
200 42551C N(4)-acetylcytidine-5'-monophosphate pac4C nucleotide None 365.233 not aplicable Natural No Yes Cytidine

4AC

81
102 02G N2,2'-O-dimethylguanosine m2Gm γ nucleoside C12O5N5H17 311.123 311.294 not applicable Natural No Yes Guanosine
142 027G N2,7,2'-O-trimethylguanosine m2,7Gm æ nucleoside C13H21N5O5 327.1543 326.328 not applicable Natural No Yes Guanosine
4 27G N2,7-dimethylguanosine m2,7G nucleoside C12O5N5H19 313.1386 312.302 not applicable Natural No Yes Guanosine 133
133 279553N N2,7-dimethylguanosine cap (cap DMG) m2,7GpppN ® nucleotide C12H17N5O11P2 469.04 664.325 unspecified residue Natural No Yes Adenosine, Uridine, Cytidine, Guanosine
40 022G N2,N2,2'-O-trimethylguanosine m2,2Gm | nucleoside C13O5N5H19 325.1386 325.32 not applicable Natural No Yes Guanosine
111 227G N2,N2,7-trimethylguanosine m2,2,7G nucleoside C13O5N5H21 327.1543 326.328 not applicable Natural No Yes Guanosine 134
134 2279553N N2,N2,7-trimethylguanosine cap (cap TMG) m2,2,7GpppN nucleotide C13H19N5O11P2 483.0556 678.352 unspecified residue Natural No Yes Adenosine, Uridine, Cytidine, Guanosine
13 22G N2,N2-dimethylguanosine m2,2G R nucleoside C12O5N5H17 311.123 311.294 not applicable Natural No Yes Guanosine 198
198 22551G N2-dimethylguanosine-5'-monophosphate pm2,2G nucleotide None 391.274 not aplicable Natural No Yes Guanosine

M2G

13
36 2G N2-methylguanosine m2G L nucleoside C11O5N5H15 297.1073 297.267 not applicable Natural No Yes Guanosine 191
39 04C N4,2'-O-dimethylcytidine m4Cm λ nucleoside C11O5N3H17 271.1168 271.27 not applicable Natural No Yes Cytidine 208
69 044C N4,N4,2'-O-trimethylcytidine m4,4Cm β nucleoside C12O5N3H19 285.1325 285.296 not applicable Natural No Yes Cytidine
15 44C N4,N4-dimethylcytidine m4,4C μ nucleoside C11O5N3H17 271.1168 271.27 not applicable Natural No Yes Cytidine 250
106 042C N4-acetyl-2'-O-methylcytidine ac4Cm nucleoside C12O6N3H17 299.1117 299.28 not applicable Natural No Yes Cytidine 257
81 42C N4-acetylcytidine ac4C M nucleoside C11O6N3H15 285.0961 285.253 not applicable Natural No Yes Cytidine 200
9 4C N4-methylcytidine m4C ν nucleoside C10O5N3H15 257.1012 257.243 not applicable Natural No Yes Cytidine 281
82 06A N6,2'-O-dimethyladenosine m6Am χ nucleoside C12O4N5H17 295.1281 295.294 not applicable Natural No Yes Adenosine
124 066A N6,N6,2'-O-trimethyladenosine m6,6Am η nucleoside C13O4N5H19 309.1437 309.321 not applicable Natural No Yes Adenosine
41 66A N6,N6-dimethyladenosine m6,6A ζ nucleoside C12O4N5H17 295.1281 295.294 not applicable Natural No Yes Adenosine 192
10 60A N6-(cis-hydroxyisopentenyl)adenosine io6A ` nucleoside C15O5N5H21 351.1543 351.358 not applicable Natural No Yes Adenosine
125 64A N6-acetyladenosine ac6A nucleoside C12O5N5H15 309.1073 309.278 not applicable Natural No Yes Adenosine
177 67A N6-formyladenosine f6A Ϩ nucleoside C11H13N5O5 295.0917 295.251 not applicable Natural No Yes Adenosine
114 65A N6-glycinylcarbamoyladenosine g6A nucleoside C13O7N6H16 368.108 368.302 not applicable Natural No Yes Adenosine
176 68A N6-hydroxymethyladenosine hm6A Ϫ nucleoside C11H15N5O5 297.1073 297.267 not applicable Natural No Yes Adenosine
19 63A N6-hydroxynorvalylcarbamoyladenosine hn6A nucleoside C16O8N6H22 426.1499 426.381 not applicable Natural No Yes Adenosine
253 61551A N6-isopentenyl-adenosine-5'-monophosphate pi6A nucleotide None 417.354 not aplicable Natural No Yes Adenosine

6IA

51
56 61A N6-isopentenyladenosine i6A + nucleoside C15O4N5H21 335.1594 335.358 not applicable Natural No Yes Adenosine 253
16 662A N6-methyl-N6-threonylcarbamoyladenosine m6t6A E nucleoside C16O8N6H22 426.1499 426.381 not applicable Natural No Yes Adenosine 231
302 662551A N6-methyl-N6-threonylcarbamoyladenosine-5'-monophosphate pm6t6A nucleotide None 506.361 not aplicable Natural No Yes Adenosine

AET

130
338 64555N N6-methyl-adenosine pentaphosphate 5' cap (cap Ap5N) m6ApppppN nucleotide None None Natural No Yes Adenosine, Cytidine, Guanosine, Uridine
337 64554N N6-methyl-adenosine tetraphosphate 5' cap (cap Ap4N) m6AppppN nucleotide None None Natural No Yes Adenosine, Cytidine, Guanosine, Uridine
336 64553N N6-methyl-adenosine triphosphate 5' cap (cap A) m6ApppN nucleotide None None Natural No Yes Adenosine, Cytidine, Guanosine, Uridine
96 6A N6-methyladenosine m6A = nucleoside C11O4N5H15 281.1124 281.268 not applicable Natural No Yes Adenosine 211
211 6551A N6-methyladenosine-5'-monophosphate pm6A nucleotide None 361.248 not aplicable Natural No Yes Adenosine

6MZ

96
130 62A N6-threonylcarbamoyladenosine t6A 6 nucleoside C15O8N6H20 412.1343 411.37 not applicable Natural No Yes Adenosine
231 62551A N6-threonylcarbamoyladenosine-5'-monophosphate pt6A nucleotide None 492.335 not aplicable Natural No Yes Adenosine

T6A

130
339 79554N N7-methyl-guanosine tetraphosphate 5' cap (cap m7Gp4N) m7GppppN nucleotide None None Natural No Yes Adenosine, Cytidine, Guanosine, Uridine
132 79553N N7-methyl-guanosine cap (cap 0) m7GpppN © nucleotide C11H15N5O11P2 455.0243 650.298 unspecified residue Natural No Yes Adenosine, Cytidine, Guanosine, Uridine
212 7551G N7-methyl-guanosine-5'-monophosphate pm7G nucleotide None 378.255 not aplicable Natural No Yes Guanosine

G7M

116
201 0551C O2'-methylcytidine-5'-monophosphate pCm nucleotide None 337.223 not aplicable Natural No Yes Cytidine

OMC

84
193 0551G O2'-methylguanosine-5'-monophosphate pGm nucleotide None 377.247 not aplicable Natural No Yes Guanosine

OMG

28
205 0551U O2'-methyluridine 5'-monophosphate pUm nucleotide None 338.208 not aplicable Natural No Yes Uridine

OMU

50
257 042551C [(2~{R},3~{R},4~{R},5~{R})-5-(4-acetamido-2-oxidanylidene-pyrimidin-1-yl)-4-methoxy-3-oxidanyl-oxolan-2-yl]methyl dihydrogen phosphate pac4Cm nucleotide None 379.26 not aplicable Natural No Yes Cytidine

LHH

106
250 44551C [(2~{R},3~{S},4~{R},5~{R})-5-[4-(dimethylamino)-2-oxidanylidene-pyrimidin-1-yl]-3,4-bis(oxidanyl)oxolan-2-yl]methyl dihydrogen phosphate pm4,4C nucleotide None 351.25 not aplicable Natural No Yes Cytidine

LV2

15
109 A adenosine A A nucleoside C10O4N5H13 267.0968 267.241 not applicable Natural No Yes Adenosine 310
335 4555N adenosine pentaphosphate 5' cap (cap Ap5N) ApppppN nucleotide None None Natural No Yes Adenosine, Cytidine, Guanosine, Uridine
334 4554N adenosine tetraphosphate 5' cap (cap Ap4N) AppppN nucleotide None None Natural No Yes Adenosine, Cytidine, Guanosine, Uridine
333 4553N adenosine triphosphate 5' cap (cap A) ApppN nucleotide None None Natural No Yes Adenosine, Cytidine, Guanosine, Uridine
310 33551A adenosine-3',5'-diphosphate pAp nucleotide None 427.201 not aplicable Natural No Yes Adenosine

A3P

109
317 552A adenosine-5'-diphosphate nucleotide None 427.201 not aplicable Natural No Yes Adenosine 109
215 3377A adenosine-5'-phosphate-2',3'-cyclic phosphate pA2'3'cp nucleotide None 409.186 not aplicable Natural No Yes Adenosine

A23

109
233 553A adenosine-5'-triphosphate pppA nucleotide None 507.181 not aplicable Natural No Yes Adenosine

ATP

109
141 20C agmatidine C+ ¿ nucleoside C14H25N7O4 355.1968 355.393 not applicable Natural No Yes Cytidine
170 2551N alpha-dimethylmonophosphate 5' cap mmpN nucleotide C7H14O10P2 320.0062 241.156 unspecified residue Natural No Yes Adenosine, Cytidine, Guanosine, Uridine
171 1551N alpha-methylmonophosphate 5' cap mpN nucleotide C6H12O10P2 305.9906 226.121 unspecified residue Natural No Yes Adenosine, Cytidine, Guanosine, Uridine
99 103G archaeosine G+ ( nucleoside C12H16N6O5 324.1182 324.293 not applicable Natural No Yes Guanosine
175 69A cyclic N6-threonylcarbamoyladenosine ct6A e nucleoside C15O7N6H18 394.1237 394.339 not applicable Natural No Yes Adenosine
46 C cytidine C C nucleoside C9O5N3H13 243.0855 243.217 not applicable Natural No Yes Cytidine 300
300 551C cytidine-5'-monophosphate pC nucleotide None 323.197 not aplicable Natural No Yes Cytidine

C5P

46
209 3377C cytidine-5'-phosphate-2',3'-cyclic phosphate pC2'3'cp nucleotide None 385.161 not aplicable Natural No Yes Cytidine

CCC

46
14 8U dihydrouridine D D nucleoside C9O6N2H14 246.0852 246.217 not applicable Natural No Yes Uridine 190
6 102G epoxyqueuosine oQ ς nucleoside C17O8N5H23 425.1547 425.393 not applicable Natural No Yes Guanosine
86 104G galactosyl-queuosine galQ 9 nucleoside C23O12N5H33 571.2126 571.534 not applicable Natural No Yes Guanosine
135 1553N gamma-methyltriphosphate 5' cap mpppN § nucleotide CH3O6P2 172.9405 384.065 unspecified residue Natural No Yes Adenosine, Cytidine, Guanosine, Uridine
42 105G glutamyl-queuosine gluQ nucleoside C22O10N6H30 538.2023 538.508 not applicable Natural No Yes Guanosine
238 3377G guanoside-5'-phosphate-2',3'-cyclic phosphate pG2'3'cp nucleotide None 425.185 not aplicable Natural No Yes Guanosine

23G

38
38 G guanosine G G nucleoside C10O5N5H13 283.0917 283.241 not applicable Natural No Yes Guanosine 235
161 551G551N guanosine added to any ribonucleotide pG(pN) none nucleotide C10H13N5O8P 362.0502 555.284 unspecified residue Natural No Yes Adenosine, Uridine, Cytidine, Guanosine
148 9553N guanosine triphosphate 5' cap (cap G) GpppN ϑ nucleotide C10H12N5O14P3 440.0008 635.264 unspecified residue Natural No Yes Adenosine, Cytidine, Guanosine, Uridine
259 33551G guanosine-3',5'-diphosphate pGp nucleotide None 443.201 not aplicable Natural No Yes Guanosine

PGP

38
218 552G guanosine-5'-diphosphate ppG nucleotide None 443.201 not aplicable Natural No Yes Guanosine

GDP

38
235 551G guanosine-5'-monophosphate pG nucleotide None 283.241 not aplicable Natural No Yes Guanosine

5GP

38
202 553G guanosine-5'-triphosphate pppG nucleotide None 523.18 not aplicable Natural No Yes Guanosine

GTP

38
181 2165A hydroxy-N6-threonylcarbamoyladenosine ht6A « nucleoside C15H20N6O9 428.1292 428.354 not applicable Natural No Yes Adenosine
97 34830G hydroxywybutosine OHyW nucleoside C21O10N6H28 524.1867 524.481 not applicable Natural No Yes Guanosine
113 9A inosine I I nucleoside C10O5N4H12 268.0808 268.226 not applicable Natural No Yes Adenosine 217
217 9551A inosine-5'-monophosphate pI nucleotide None 348.206 not aplicable Natural No Yes Adenosine

I

113
89 42G isowyosine imG2 nucleoside C14O5N5H17 335.123 335.315 not applicable Natural No Yes Guanosine
90 106G mannosyl-queuosine manQ 8 nucleoside C23O12N5H33 571.2126 571.534 not applicable Natural No Yes Guanosine
160 3480G methylated undermodified hydroxywybutosine OHyWy y nucleoside C19H26N6O8 466.1812 466.445 not applicable Natural No Yes Guanosine
59 342G methylwyosine mimG nucleoside C15O5N5H19 349.1386 349.342 not applicable Natural No Yes Guanosine
121 34832G peroxywybutosine o2yW W nucleoside C21O11N6H28 540.1816 540.481 not applicable Natural No Yes Guanosine
118 9U pseudouridine Y P nucleoside C9O6N2H12 244.0695 244.201 not applicable Natural No Yes Uridine 185
185 9551U pseudouridine-5'-monophosphate pY nucleotide None 324.181 not aplicable Natural No Yes Uridine

PSU

118
105 10000G queuine Qbase base C12O3N5H15 277.1175 277.279 not applicable Natural No Yes Guanosine 65
65 10G queuosine Q Q nucleoside C17O7N5H23 409.1597 409.394 not applicable Natural No Yes Guanosine
93 3470G undermodified hydroxywybutosine OHyWx š nucleoside C18O8N6H24 452.1655 452.419 not applicable Natural No Yes Guanosine
165 551N unknown 5' monophosphate ribonucleotide pN m nucleotide C5H7O10P2 288.9514 211.087 unspecified residue Natural No Yes Adenosine, Uridine, Cytidine, Guanosine
98 X unknown modification xX @ nucleoside - 0.0 0.0 unknown residue, modification present but its character is unknown Natural No Yes Adenosine, Uridine, Cytidine, Guanosine
72 ?A unknown modified adenosine xA H nucleoside - 0.0 0.0 unknown modification Natural No Yes Adenosine
70 ?C unknown modified cytidine xC < nucleoside - 0.0 0.0 unknown modification Natural No Yes Cytidine
66 ?G unknown modified guanosine xG ; nucleoside - 0.0 0.0 unknown modification Natural No Yes Guanosine
76 ?U unknown modified uridine xU N nucleoside - 0.0 0.0 unknown modification Natural No Yes Uridine
25 0X unknown nucleoside 2'-O-methylated Xm Î nucleoside - 0.0 0.0 unknown base, ribose 2'-O-methylated Natural No Yes Adenosine, Uridine, Cytidine, Guanosine
140 N unknown ribonucleoside residue N . nucleoside - 0.0 0.0 unspecified residue Natural No Yes Adenosine, Uridine, Cytidine, Guanosine
60 U uridine U U nucleoside C9O6N2H12 244.0695 244.201 not applicable Natural No Yes Uridine 255
47 502U uridine 5-oxyacetic acid cmo5U V nucleoside C11O9N2H14 318.0699 318.237 not applicable Natural No Yes Uridine 241
58 503U uridine 5-oxyacetic acid methyl ester mcmo5U υ nucleoside C12O9N2H16 332.0856 332.263 not applicable Natural No Yes Uridine
255 551U uridine-5'-monophosphate pU nucleotide None 324.181 not aplicable Natural No Yes Uridine

U5P

60
268 3377551U uridine-5'-monophosphate-2',3'-cyclic phosphate pU2'3'cp nucleotide None 386.146 not aplicable Natural No Yes Uridine

U23

60
30 3483G wybutosine yW Y nucleoside C21O9N6H28 508.1918 508.482 not applicable Natural No Yes Guanosine 244
213 3483551G wybutosine-5'-monophosphate pyW nucleotide None 588.462 not aplicable Natural No Yes Guanosine

YG

30
244 34831551G wybutosine[C15(S)]-5'-monophosphate pyW nucleotide None 588.462 not aplicable Natural No Yes Guanosine

YYG

30
139 34G wyosine imG nucleoside C14O5N5H17 335.123 335.315 not applicable Natural No Yes Guanosine
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